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Future projections of southwestern African hydroclimate are highly uncertain. However, insights from past warm climates, like the Pliocene, can reveal mechanisms of future change and help benchmark models. Using leaf wax hydrogen isotopes to reconstruct precipitation (δD_p) from Namibia over the past 5 million years, we find a long‐term depletion trend (−50‰). Empirical mode decomposition indicates this trend is linked to sea surface temperatures (SSTs) within the Benguela Upwelling System, but modulated by Indian Ocean SSTs on shorter timescales. The influence of SSTs on reconstructed regional hydroclimate is similar to that observed during modern Benguela Nio events, which bring extreme flooding to the region. Isotope‐enabled simulations and PlioMIP2 results suggest that capturing a Benguela Nio‐like state is key to accurately simulating Pliocene, and future, regional hydroclimate. This has implications for future regional climate, since an increased frequency of Benguela Nios poses risk to the ecosystems and industries in the region.

The Pliocene offers insights into future climate, with near‐modern atmospheric pCO₂and global mean surface temperature estimated to be 3–4°C above pre‐industrial. However, the hydrological response differs between future global warming and early Pliocene climate model simulations. This discrepancy results from the use of reduced meridional and zonal sea surface temperature (SST) gradients, based on foraminiferal Mg/Ca and Alkenone proxy evidence, to force the early Pliocene simulation. Subsequent, SST reconstructions based on the organic proxy TEX₈₆, have found warmer temperatures in the warm pool, bringing the magnitude of the gradient reductions into dispute. We design an independent test of Pliocene SST scenarios and their hydrological cycle “fingerprints.” We use an isotope‐enabled General Circulation Model, iCAM5, to model the distribution of water isotopes in precipitation in response to four climatological SST and sea‐ice fields representing modern, abrupt 4 × CO₂, late Pliocene and early Pliocene climates. We conduct a proxy‐model comparison with all the available precipitation isotope proxy data, and we identify target regions that carry precipitation isotopic fingerprints of SST gradients as priorities for additional proxy reconstructions. We identify two regions with distinct precipitation isotope (D/H) fingerprints resulting from reduced SST gradients: the Maritime Continent (D‐enriched due to reduced convective rainfall) and the Sahel (wetter, more deep convection, D‐depleted). The proxy‐model comparison using available plant wax reconstructions, mostly from Africa, is promising but inconclusive. Additional proxy reconstructions are needed in both target regions and in much of the world for significant tests of SST scenarios and dynamical linkages to the hydrological cycle.

Earth's hydrological cycle is expected to intensify in response to global warming, with a “wet‐gets‐wetter, dry‐gets‐drier” response anticipated over the ocean. Subtropical regions (∼15°–30°N/S) are predicted to become drier, yet proxy evidence from past warm climates suggests these regions may be characterized by wetter conditions. Here we use an integrated data‐modeling approach to reconstruct global and zonal‐mean rainfall patterns during the early Eocene (∼56–48 million years ago). The Deep‐Time Model Intercomparison Project (DeepMIP) model ensemble indicates that the mid‐ (30°–60°N/S) and high‐latitudes (>60°N/S) are characterized by a thermodynamically dominated hydrological response to warming and overall wetter conditions. The tropical band (0°–15°N/S) is also characterized by wetter conditions, with several DeepMIP models simulating narrowing of the Inter‐Tropical Convergence Zone. However, the latter is not evident from the proxy data. The subtropics are characterized by negative precipitation‐evaporation anomalies (i.e., drier conditions) in the DeepMIP models, but there is surprisingly large inter‐model variability in mean annual precipitation (MAP). Intriguingly, we find that models with weaker meridional temperature gradients (e.g., CESM, GFDL) are characterized by a reduction in subtropical moisture divergence, leading to an increase in MAP. These model simulations agree more closely with our new proxy‐derived precipitation reconstructions and other key climate metrics and imply that the early Eocene was characterized by reduced subtropical moisture divergence. If the meridional temperature gradient was even weaker than suggested by those DeepMIP models, circulation‐induced changes may have outcompeted thermodynamic changes, leading to wetter subtropics. This highlights the importance of accurately reconstructing zonal temperature gradients when reconstructing past rainfall patterns.